How does the startup code work?

I would like to understand how the code works when one first goes to SugarCRM.

I am upgrading from 8.0.2 to 9.0.3.

I upgraded our TEST environment.  Then upgrade our PROD environment.  After upgrading PROD the changes were committed to GITHUB.  When I pulled the changes committed to GITHUB from PROD, I get the loading... screen when I am trying to access TEST.

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